Subject of the sentence is "Delta, United, and American". This is a plural subject. Each after the plural subject will have a plural verb.
Each before the plural subject will have a singular verb.
Eg: Winship estimates thateach of the three largest carriers - Delta, United, and American - generates roughly $1billion annually from the sale of frequent-flier miles to hotels, rental-car companies, and others offering points-for-miles to their customers
...
Each before the plural subject will have a singular verb.
Eg: Winship estimates thateach of the three largest carriers - Delta, United, and American - generates roughly $1billion annually from the sale of frequent-flier miles to hotels, rental-car companies, and others offering points-for-miles to their customers
...
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