Art experts have attributed the enduring intrigue of Leonardo da Vinci's Mona Lisa to everything from having an enigmatic smile, which was never explained, to her association with the rich and powerful families of Europe.
• having an enigmatic smile, which was never explained- Missing Parallelish from x to y - also which was never explained is incorrect - should be present continuous tense
• having an enigmatic smile, which has never been explained- Missing Parallelish from x to y
• her
...
• having an enigmatic smile, which was never explained- Missing Parallelish from x to y - also which was never explained is incorrect - should be present continuous tense
• having an enigmatic smile, which has never been explained- Missing Parallelish from x to y
• her
...
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